Physiology · Endocrine Physiology (Pituitary, Thyroid, Adrenal, Pancreas)

Thyroid hormone (T3) exerts genomic effects by binding to thyroid hormone receptors (TRs). In the absence of T3, unliganded TR bound to thyroid response elements (TREs) on target gene promoters:

  • A Has no effect on transcription and dissociates from DNA until T3 is present
  • B Actively represses transcription by recruiting histone deacetylase (HDAC)-containing co-repressor complexes
  • C Constitutively activates basal transcription independently of T3 binding
  • D Promotes methylation of CpG islands in promoter regions, causing irreversible silencing
Correct answer: B. Actively represses transcription by recruiting histone deacetylase (HDAC)-containing co-repressor complexes

Explanation

Unliganded TR (particularly TRbeta1) binds TREs and recruits co-repressor complexes including NCoR and SMRT, which associate with histone deacetylases. This creates a transcriptionally repressed chromatin state. On T3 binding, the co-repressor complex is replaced by co-activator complexes (SRC/p160 family) with histone acetyltransferase activity, opening chromatin and activating transcription. This explains why hypothyroidism is not merely reduced T3 action but an actively repressive state — accounting for the disproportionate severity of congenital hypothyroidism.

Reference: Guyton & Hall, Textbook of Medical Physiology, 14th ed.

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